J. T. Beatty, K. Bendschneider, and R. J. Robinson, On the natural selection and evolution of the aerobic phototrophic bacteria A new spectrophotometric determination of nitrite in seawater, Photosynthesis Research, vol.73, issue.1/3, pp.109-114, 1952.
DOI : 10.1023/A:1020493518379

R. Benner and M. Strom, A critical evaluation of the analytical blank associated with DOC measurements by high-temperature catalytic oxidation, Marine Chemistry, vol.41, issue.1-3, pp.153-160, 1993.
DOI : 10.1016/0304-4203(93)90113-3

T. Berman, P. D. Walline, A. Schneller, J. Rothenberg, T. et al., Secchi disk depth record: A claim for the eastern Mediterranean1, Limnology and Oceanography, vol.30, issue.2, pp.447-448, 1985.
DOI : 10.4319/lo.1985.30.2.0447

J. C. Cho, M. D. Stapels, R. M. Morris, K. L. Vergin, M. S. Schwalbach et al., Polyphyletic photosynthetic reaction centre genes in oligotrophic marine Gammaproteobacteria, Polyphyletic photosynthetic reaction centre genes in oligotrophic marine Gammaproteobacteria, pp.1456-1463, 2007.
DOI : 10.1126/science.1093857

M. T. Cottrell and D. L. Kirchman, Photoheterotrophic Microbes in the Arctic Ocean in Summer and Winter, Applied and Environmental Microbiology, vol.75, issue.15, pp.4958-4966, 2009.
DOI : 10.1128/AEM.00117-09

M. T. Cottrell, A. Mannino, and D. L. Kirchman, Aerobic Anoxygenic Phototrophic Bacteria in the Mid-Atlantic Bight and the North Pacific Gyre, Applied and Environmental Microbiology, vol.72, issue.1, pp.557-564, 2006.
DOI : 10.1128/AEM.72.1.557-564.2006

M. T. Cottrell, J. Ras, and D. L. Kirchman, Bacteriochlorophyll and community structure of aerobic anoxygenic phototrophic bacteria in a particle-rich estuary, The ISME Journal, vol.6, issue.7, pp.945-954, 2010.
DOI : 10.1111/j.1462-2920.2007.01265.x

Y. Crombet, K. Leblanc, B. Quéguiner, T. Moutin, P. Rimmelin et al., Deep silicon maxima in the stratified oligotrophic Mediterranean Sea, Biogeosciences, vol.85194, pp.459-47510, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00700258

D. De-corte, T. Yokokawa, M. M. Varela, H. Agogue, and G. J. Herndl, Spatial distribution of Bacteria and Archaea and amoA gene copy numbers throughout the water column of the Eastern Mediterranean Sea, The ISME Journal, vol.24, issue.2, pp.147-158, 2009.
DOI : 10.1038/ismej.2008.94

A. Eiler, Evidence for the Ubiquity of Mixotrophic Bacteria in the Upper Ocean: Implications and Consequences, Applied and Environmental Microbiology, vol.72, issue.12, pp.7431-7437, 2006.
DOI : 10.1128/AEM.01559-06

J. M. Gonzalez, R. Simo, R. Massana, J. S. Covert, E. O. Casamayor et al., Bacterial Community Structure Associated with a Dimethylsulfoniopropionate-Producing North Atlantic Algal Bloom, Applied and Environmental Microbiology, vol.66, issue.10, pp.4237-4246, 2000.
DOI : 10.1128/AEM.66.10.4237-4246.2000

M. R. Holmes, A. Aminot, R. Kérouel, B. A. Hooker, and B. J. Peterson, A simple and precise method for measuring ammonium in marine and freshwater ecosystems, Canadian Journal of Fisheries and Aquatic Sciences, vol.56, issue.10, pp.1801-1808, 1999.
DOI : 10.1139/f99-128

L. Ignatiades, Scaling the trophic status of the Aegean Sea, eastern Mediterranean, Journal of Sea Research, vol.54, issue.1, pp.51-57, 2005.
DOI : 10.1016/j.seares.2005.02.010

C. Jeanthon, D. Boeuf, O. Dahan, L. Gall, F. Garczarek et al., Diversity of cultivated and metabolically active aerobic anoxygenic phototrophic bacteria along an oligotrophic gradient in the Mediterranean Sea, Biogeosciences Discuss, 2011.
URL : https://hal.archives-ouvertes.fr/hal-01218510

N. Jiao, Y. Zhang, Y. H. Zeng, N. Hong, R. Liu et al., Distinct distribution pattern of abundance and diversity of aerobic anoxygenic phototrophic bacteria in the global ocean, Environmental Microbiology, vol.62, issue.12, pp.3091-3099, 2007.
DOI : 10.1360/03wd0156

D. M. Karl, Microbiological oceanography: Hidden in a sea of microbes, Nature, vol.415, issue.6872, pp.590-591, 2002.
DOI : 10.1038/415590b

D. M. Karl and G. Tien, MAGIC: A sensitive and precise method for measuring dissolved phosphorus in aquatic environments, Limnology and Oceanography, vol.37, issue.1, pp.105-116, 1992.
DOI : 10.4319/lo.1992.37.1.0105

M. Koblí?ek, O. Béjà, R. R. Bidigare, S. Christensen, B. Benitez-nelson et al., Isolation and characterization of Erythrobacter sp. strains from the upper ocean, Archives of Microbiology, vol.68, issue.5, pp.327-338, 2003.
DOI : 10.1007/s00203-003-0596-6

M. Koblí?ek, J. Sto´nsto´n-egiert, S. Sagan, and Z. S. Kolber, Diel changes in bacteriochlorophyll a concentration suggest rapid bacterioplankton cycling in the Baltic Sea, FEMS Microbiol. Ecol, vol.51, pp.353-361, 2005.

M. Koblí?ek, P. G. Falkowski, and Z. S. Kolber, Diversity and distribution of photosynthetic bacteria in the Black Sea, Deep- Sea Res, Pt. II, vol.53, pp.1934-1944, 2006.

M. Koblí?ek, M. Ma?ín, J. Ras, A. J. Poulton, and O. Prá?il, Rapid growth rates of aerobic anoxygenic phototrophs in the ocean, Environmental Microbiology, vol.49, issue.10, pp.2401-2406, 2007.
DOI : 10.1007/BF00244268

Z. S. Kolber, C. L. Van-dover, R. A. Niederman, and P. G. Falkowski, Bacterial photosynthesis in surface waters of the open ocean, Nature, vol.407, pp.177-179, 2000.

Z. S. Kolber, F. G. Plumley, A. S. Lang, J. T. Beatty, R. E. Blankenship et al., Contribution of Aerobic Photoheterotrophic Bacteria to the Carbon Cycle in the Ocean, Science, vol.292, issue.5526, pp.2492-2495, 2001.
DOI : 10.1126/science.1059707

D. Lamy, D. Carvalho-maalouf, P. Cottrell, M. T. Lami, R. Catala et al., Seasonal dynamics of aerobic anoxygenic phototrophs in a Mediterranean coastal lagoon, Seasonal dynamics of aerobic phototrophs in a Mediterranean coastal lagoon, pp.153-163, 2011.
DOI : 10.3354/ame01467

R. Lami, M. T. Cottrell, J. Ras, O. Ulloa, I. Obernosterer et al., High Abundances of Aerobic Anoxygenic Photosynthetic Bacteria in the South Pacific Ocean, High abundances of aerobic anoxygenic photosynthetic bacteria in the South Pacific Ocean, pp.4198-4205, 2007.
DOI : 10.1128/AEM.02652-06

R. Lami, Z. Euperová, J. Ras, P. Lebaron, M. Koblí?ek et al., Distribution of free-living and particle-attached aerobic anoxygenic phototrophic bacteria in marine environments, Aquatic Microbial Ecology, vol.55, issue.74, pp.31-38, 2009.
DOI : 10.3354/ame01282

D. C. López-sandoval, A. Fernández, and E. Marañón, Dissolved and particulate primary production along a longitudinal www.biogeosciences.net, Biogeosciences, vol.8973, issue.8, pp.973-985, 2011.

D. Lamy, Aerobic anoxygenic phototrophs in the Mediterranean Sea gradient in the Mediterranean Sea, Biogeosciences, vol.85194, issue.10, pp.815-825, 2011.

D. Marie, N. Simon, L. Guillou, F. Partensky, D. Vaulot et al., Flow cytometric analysis of marine picoplankton, Protocols in flow cytometry and cell sorting, pp.421-454, 2002.

M. Ma?ín, J. Nedoma, L. Pechar, and M. Koblí?ek, Distribution of aerobic anoxygenic phototrophs in temperate freshwater systems, Environmental Microbiology, vol.9, issue.8, 1988.
DOI : 10.1111/j.1462-2920.2008.01615.x

M. A. Moran and W. L. Miller, Resourceful heterotrophs make the most of light in the coastal ocean, Nature Reviews Microbiology, vol.444, issue.10, pp.792-800, 2007.
DOI : 10.1038/nrmicro1746

T. Moutin and P. Raimbault, Primary production, carbon export and nutrients availability in western and eastern Mediterranean Sea in early summer, J. Marine Syst, pp.33-34, 1996.

T. Moutin, T. F. Thingstad, F. Van-wambeke, D. Marie, G. Slawyk et al., ?, Limnology and Oceanography, vol.47, issue.5, pp.1562-1567, 2002.
DOI : 10.4319/lo.2002.47.5.1562

T. Moutin, D. M. Karl, S. Duhamel, P. Rimmelin, P. Raimbault et al., Phosphate availability and the ultimate control of new nitrogen input by nitrogen fixation in the tropical Pacific Ocean, Biogeosciences, vol.55194, pp.95-10910, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00330253

T. Moutin, F. Van-wambeke, and L. Prieur, Introduction to the Biogeochemistry from the Oligotrophic to the Ultraoligotrophic Mediterranean (BOUM) experiment, Biogeosciences Discuss, 2011.

J. Murphy and J. P. Riley, A modified single solution method for the determination of phosphate in natural waters, Analytica Chimica Acta, vol.27, pp.31-36, 1962.
DOI : 10.1016/S0003-2670(00)88444-5

A. Oz, G. Sabehi, M. Koblí?ek, R. Massana, and O. Béjà, Roseobacter-Like Bacteria in Red and Mediterranean Sea Aerobic Anoxygenic Photosynthetic Populations, Applied and Environmental Microbiology, vol.71, issue.1, pp.344-353, 2005.
DOI : 10.1128/AEM.71.1.344-353.2005

J. Pinhassi, L. Gomez-consarnau, L. Alonso-saez, M. M. Sala, M. Vidal et al., Seasonal changes in bacterioplankton nutrient limitation and their effects on bacterial community composition in the NW Mediterranean Sea, Aquatic Microbial Ecology, vol.44, pp.241-252, 2006.
DOI : 10.3354/ame044241

M. Pujo-pay and P. Raimbault, improvement of the wet-oxidation procedure for simultaneous determination of particulate organic nitrogen and phosphorus collected on filters, Marine Ecology Progress Series, vol.105, pp.203-207, 1994.
DOI : 10.3354/meps105203

M. Pujo-pay, P. Conan, R. , and P. , Excretion of dissolved organic nitrogen by phytoplankton assessed by wet oxidation and 15N tracer procedures, Marine Ecology Progress Series, vol.153, pp.99-111, 1997.
DOI : 10.3354/meps153099

M. Pujo-pay, P. Conan, L. Oriol, V. Cornet-barthaux, C. Falco et al., Integrated survey of elemental stoichiometry (C, N, P) from the western to eastern Mediterranean Sea, Biogeosciences, vol.85194, pp.883-89910, 2011.

A. Ramette, Multivariate analyses in microbial ecology, FEMS Microbiology Ecology, vol.62, issue.2, pp.142-160, 2007.
DOI : 10.1111/j.1574-6941.2007.00375.x

J. Ras, H. Claustre, and J. Uitz, Spatial variability of phytoplankton pigment distributions in the Subtropical South Pacific Ocean: comparison between in situ and predicted data, Biogeosciences, vol.5, issue.2, pp.353-36910, 2008.
DOI : 10.5194/bg-5-353-2008

URL : https://hal.archives-ouvertes.fr/hal-00330697

P. Rimmelin and T. Moutin, Re-examination of the MAGIC method to determine low orthophosphate concentration in seawater, Analytica Chimica Acta, vol.548, issue.1-2, pp.174-182, 2005.
DOI : 10.1016/j.aca.2005.05.071

I. Salka, V. Moulisová, M. Koblí?ek, G. Jost, K. Jürgens et al., Abundance, Depth Distribution, and Composition of Aerobic Bacteriochlorophyll a-Producing Bacteria in Four Basins of the Central Baltic Sea, Applied and Environmental Microbiology, vol.74, issue.14, pp.4398-4404, 2008.
DOI : 10.1128/AEM.02447-07

M. S. Schwalbach and J. A. Fuhrman, Wide-ranging abundances of aerobic anoxygenic phototrophic bacteria in the world ocean revealed by epifluorescence microscopy and quantitative PCR, Limnology and Oceanography, vol.50, issue.2, pp.620-628, 2005.
DOI : 10.4319/lo.2005.50.2.0620

J. H. Sharp, Improved analysis for ???particulate??? organic carbon and nitrogen from seawater1, Limnology and Oceanography, vol.19, issue.6, pp.984-989, 1974.
DOI : 10.4319/lo.1974.19.6.0984

T. Shiba, Roseobacter litoralis gen. nov., sp. nov., and Roseobacter denitrificans sp. nov., Aerobic Pink-Pigmented Bacteria which Contain Bacteriochlorophyll a, Systematic and Applied Microbiology, vol.14, issue.2, pp.140-145, 1991.
DOI : 10.1016/S0723-2020(11)80292-4

M. E. Sieracki, I. C. Gilg, E. C. Thier, N. J. Poulton, and R. Goericke, Distribution of planktonic aerobic anoxygenic photoheterotrophic bacteria in the northwest Atlantic, Limnology and Oceanography, vol.51, issue.1, pp.38-46, 2006.
DOI : 10.4319/lo.2006.51.1.0038

D. C. Smith and F. Azam, A simple and economical method for measuring bacterial protein synthesis rates in seawater using 3H- leucine, Mar. Microb. Food Webs, vol.6, pp.107-114, 1992.

M. T. Suzuki and O. Béjà, An elusive marine photosynthetic bacterium is finally unveiled, Natl. Acad. Sci. USA, pp.2561-2562, 2007.
DOI : 10.1073/pnas.0700175104

M. T. Suzuki, C. M. Preston, F. P. Chavez, and E. F. Delong, Quantitative mapping of bacterioplankton populations in seawater: field tests across an upwelling plume in Monterey Bay, Aquatic Microbial Ecology, vol.24, pp.117-127, 2001.
DOI : 10.3354/ame024117

T. Tanaka, T. F. Thingstad, U. Christaki, J. Colombet, V. Cornet-barthaux et al., Lack of P-limitation of phytoplankton and heterotrophic prokaryotes in surface waters of three anticyclonic eddies in the stratified Mediterranean Sea, Biogeosciences, vol.8, issue.2, pp.525-53810, 2011.
DOI : 10.5194/bg-8-525-2011

URL : https://hal.archives-ouvertes.fr/hal-00573163

C. J. Ter-braak and P. Smilauer, CANOCO ??? an extension of DECORANA to analyze species-environment relationships, Hydrobiologia, vol.18, issue.3, pp.169-170, 1989.
DOI : 10.1007/BF02392953

T. F. Thingstad, U. L. Zweifel, and F. And-rassoulzadegan, P limitation of heterotrophic bacteria and phytoplankton in the northwest Mediterranean, Limnology and Oceanography, vol.43, issue.1, pp.88-94, 1998.
DOI : 10.4319/lo.1998.43.1.0088

P. Tréguer, L. Corre, and P. , Manuel d'analyse des sels nutritifs dans l'eau de mer, in: Utilisation de l'Autoanalyzer II Technicon, p.110, 1975.

L. Van-heukelem and C. S. Thomas, Computer-assisted high-performance liquid chromatography method development with applications to the isolation and analysis of phytoplankton pigments, Journal of Chromatography A, vol.910, issue.1, pp.31-49, 2001.
DOI : 10.1016/S0378-4347(00)00603-4

F. Van-wambeke, U. Christaki, A. Giannokourou, T. Moutin, and K. Souvemerzoglou, Longitudinal and Vertical Trends of Bacterial Limitation by Phosphorus and Carbon in the Mediterranean Sea, Microbial Ecology, vol.43, issue.1, pp.119-133, 2002.
DOI : 10.1007/s00248-001-0038-4

F. Van-wambeke, J. Nedoma, S. Duhamel, and P. Lebaron, Alkaline phosphatase activity of marine bacteria studied with ELF 97 substrate: success and limits in the P-limited Mediterranean Sea, Aquatic Microbial Ecology, vol.52, pp.245-251, 2008.
DOI : 10.3354/ame01238

URL : https://hal.archives-ouvertes.fr/hal-00321319

F. Van-wambeke, P. Catala, and P. Lebaron, Relationships between cytometric characteristics of high and low nucleicacid bacterioplankton cells, bacterial production and environmental parameters along a longitudinal gradient across the Mediterranean Sea, Biogeosciences Discuss, vol.75194, pp.8245-827910, 2010.

L. A. Waidner and D. L. Kirchman, Aerobic Anoxygenic Phototrophic Bacteria Attached to Particles in Turbid Waters of the Delaware and Chesapeake Estuaries, Applied and Environmental Microbiology, vol.73, issue.12, pp.3936-3944, 2007.
DOI : 10.1128/AEM.00592-07

C. Winter, M. E. Kerros, and M. G. Weinbauer, Seasonal changes of bacterial and archaeal communities in the dark ocean: Evidence from the Mediterranean Sea, Limnology and Oceanography, vol.54, issue.1, pp.160-170, 2009.
DOI : 10.4319/lo.2009.54.1.0160

E. D. Wood, F. A. Amstrong, and F. A. Richards, Determination of nitrate in sea water by cadmium-copper reduction to nitrite, Journal of the Marine Biological Association of the United Kingdom, vol.9, issue.01, pp.23-31, 1967.
DOI : 10.1016/S0003-2670(00)88614-6

V. V. Yurkov and J. T. Beatty, Aerobic anoxygenic phototrophic bacteria, Microbiol, Mol. Biol. Rev, vol.62, pp.695-724, 1998.

V. V. Yurkov and J. Csotonyi, New Light on Aerobic Anoxygenic Phototrophs, New light on aerobic anoxygenic phototrophs, pp.31-55, 2009.
DOI : 10.1007/978-1-4020-8815-5_3

N. Yutin and O. Béjà, Putative novel photosynthetic reaction centre organizations in marine aerobic anoxygenic photosynthetic bacteria: insights from metagenomics and environmental genomics, Environmental Microbiology, vol.62, issue.Pt 2, pp.2027-2033, 2005.
DOI : 10.1111/j.1462-2920.2005.00843.x

N. Yutin, M. T. Suzuki, H. Teeling, M. Weber, J. C. Venter et al., Assessing diversity and biogeography of aerobic anoxygenic phototrophic bacteria in surface waters of the Atlantic and Pacific Oceans using the Global Ocean Sampling expedition metagenomes, Environmental Microbiology, vol.62, issue.6, pp.1464-1475, 2007.
DOI : 10.1128/AEM.71.12.8958-8962.2005

N. Yutin, O. Béjà, and M. T. Suzuki, The use of denaturing gradient gel electrophoresis with fully degenerate pufM primers to monitor aerobic anoxygenic phototrophic assemblages, Limnology and Oceanography: Methods, vol.6, issue.9, pp.427-440, 2008.
DOI : 10.4319/lom.2008.6.427